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“The joy of discovery is certainly the liveliest that the mind of man can ever feel”

- Claude Bernard -

60+ peer-reviewed publications | H-index 29 | 2,370+ citations:

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63. Monitoring and modelling the dynamics of Halophila stipulacea meadows using satellite imagery and machine Learning techniques. Cohen, T.A., Rilov, G., Winters, G., Brook, A. (2026). In: Balke, WT., et al. New Trends in Theory and Practice of Digital Libraries. TPDL 2025. Communications in Computer and Information Science, vol 2694. Springer, Cham.

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62. Seagrass as a stabilizing environment for benthic foraminifera living in anthropogenically impacted coastal areas. Yahel Eshed, Gidon Winters, Gilad Antler, Sigal Abramovich, Sarit Ashckenazi-Polivoda (2025). Marine Pollution Bulletin 221: 118506.

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61. Estimation of Aboveground Biomass of Acacia Trees in the Hyper-Arid Arava, Israel Using Allometric Analysis. Gidon Winters, Casey Alexander, Thanh Hoai Tran, Giorgio Matteucci, Elli Groner (2025). Open Journal of Ecology 15(2), 194-204.

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60. Trends in seagrass research in the 21st century–are we there yet? Hung Manh Nguyen, Gidon Winters (2025). Marine Environmental Research 209: 107198.

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59. Transcriptome responses to single and combined stressors in seagrass populations from pristine and impacted sites reveal local adaptive features and core stress-response genes. Hung Manh Nguyen, Beery Yaakov, Pedro Beca-Carretero, Gabriele Procaccini, Guannan Wang, Maheshi Dassanayake, Gidon Winters, Simon Barak (2025). Science of The Total Environment 987: 179623.

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58. Helping Hand: Fungi, as Well as Bacteria, Support Ecophysiological Descriptors to Depict the Posidonia oceanica Conservation Status. Frasca, S.; Alabiso, A.; Rotini, A.; Manfra, L.; Vasquez, M.I.; Christoforou, E.; Winters, G.; Kaminer, M.; D'Andrea, M.M.; Migliore, L. (2025). Water 17: 1151.

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57. Climate Effects on Belowground Tea Litter Decomposition Depend on Ecosystem and Organic Matter Types in Global Wetlands. Stacey M. Trevathan-Tackett et al. (2025). Environmental Science & Technology.

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56. Expansion of Halophila stipulacea in parallel with declines of native seagrasses in the eastern Mediterranean Sea. Gidon Winters, Hung Manh Nguyen, Moran Kaminer (2025). Aquatic Botany 196, 103829.

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55. Strong regulation of nitrogen supply and demand in a key desert legume tree. Daphna Uni, Tamir Klein, Tania Masci, Gidon Winters, Efrat Sheffer (2024). Environmental and Experimental Botany 224, 105823.

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54. Surprising widespread Cymodocea nodosa occurrence along Israel's Mediterranean coast and Implications for Seagrass Conservation in a hotspot of climate change. Ori Hepner Ucko, Eduardo Arle, Shahar Malamud, Gidon Winters and Jonathan Belmaker (2024). Mediterranean Marine Science 25 (2), 500-510.

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53. Impacts of Desalination Brine Discharge on Benthic Ecosystems. Ryan Sirota et al. (2024). Environmental Science & Technology. 58, 13, 5631–5645.

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52. A matter of choice: Understanding the interactions between epiphytic foraminifera and their seagrass host Halophila stipulacea. Jenipher Masawa et al. (2024). Marine Environmental Research 196:106437. https://doi.org/10.1016/j.marenvres.2024.106437.

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51. Climate change and the presence of invasive species will threaten the persistence of the Mediterranean seagrass community. Pedro Beca-Carretero et al. (2024). Science of The Total Environment 910, 168675. 

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50. The effect of anaerobic remineralization of the seagrass Halophila stipulacea on porewater biogeochemistry in the Gulf of Aqaba. Neta Soto, Gidon Winters, Gilad Antler (2023). Front. Mar. Sci. 10:1250931.

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49. Undisturbed Posidonia oceanica meadows maintain the epiphytic bacterial community in different environments. Alice Rotini, Chiara Conte, Gidon Winters, Marlen I. Vasquez, Luciana Migliore (2023). Environmental Science and Pollution Research. 30, 95464–95474.

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48. Effects of anthropogenic pressures on the seagrass Halophila stipulacea and its associated macrozoobenthic communities in the northern Gulf of Aqaba. Hung Manh Nguyen, Cristina Andolina, Salvatrice Vizzini, Maria Cristina Gambi, Gidon Winters (2023). Marine Environmental Research, 189, 106073.

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47. Responses of two Acacia species to drought suggest different water-use strategies, reflecting their topographic distribution. Uni Daphna, Sheffer Efrat, Klein Tamir, Shem-Tov Rachamim, Segev Nitzan, Winters Gidon (2023). Front. Plant Sci., 14. https://doi.org/10.3389/fpls.2023.1154223

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46. Do urban tree hydraulics limit their transpirational cooling? A comparison between temperate and hot arid climates. Limor Shashua-Bar et al. (2023). Urban Climate 49, 101554. https://doi.org/10.1016/j.uclim.2023.101554

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45. Superior growth traits of invaded (Caribbean) versus native (Red Sea) populations of the seagrass Halophila stipulacea. Winters, G., Conte, C., Beca-Carretero, P. et al. (2023). Biological Invasions 25, 2325–2342. https://doi.org/10.1007/s10530-023-03045-z

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44. Gene co-expression network analysis for the selection of candidate early warning indicators of heat and nutrient stress in Posidonia oceanica. Alex Santillán-Sarmiento, Jessica Pazzaglia, Miriam Ruocco, Emanuela Dattolo, Luca Ambrosino, Gidon Winters, Lázaro Marin-Guirao, Gabriele Procaccini (2023). Science of the Total Environment, 162517. https://doi.org/10.1016/j.scitotenv.2023.162517

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43. Differential climatic conditions drive growth of Acacia tortilis tree in its range edges in Africa and Asia. Uni, D., D. Lerner, I. Smit, D. Mzimba, E. Sheffer, G. Winters, and T. Klein. (2023). American Journal of Botany e16132. https://doi.org/10.1002/ajb2.16132

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42. An invasive seagrass drives its own success in two invaded seas by both negatively affecting native seagrasses and benefiting from those costs. Chiquillo, K.L., Barber, P.H., Vasquez, M.I., Cruz-Rivera, E., Willette, D.A., Winters, G. and Fong, P. (2022). Oikos e09403. https://doi.org/10.1111/oik.09403

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41. Peak photosynthesis at summer midday in Acacia trees growing in a hyper-arid habitat. Uni Daphna, Sheffer Efrat, Winters Gidon, Lima André, Fox, Hagar and Klein Tamir. (2022). Trees. 1-13. https://doi.org/10.1007/s00468-022-02344-7.

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40. Predicted warming intensifies the negative effects of nutrient increase on tropical seagrass: A physiological and fatty acid approach. Beca-Carretero P, Azcárate-García T, Teichberg M, Patra P, Feroze F, González MJ, Medina iI, Gidon Winters (2022). Ecological Indicators 142, 109184.

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39. Editorial: Seagrasses Under Times of Change. Winters G, Teichberg M, Reuter H, Viana IG and Willette DA (2022). Front. Plant Sci. 13:87047.

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38. Teasing apart the host‑related, nutrient‑related and temperature‑related effects shaping the phenology and microbiome of the tropical seagrass Halophila stipulacea. Amir Szitenberg, Pedro Beca‑Carretero, Tomás Azcárate‑García, Timur Yergaliyev, Rivka Alexander‑Shani and Gidon Winters (2022). Environmental Microbiome 17:18.

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37. Nutrient history affects the response and resilience of the tropical seagrass Halophila stipulacea to further enrichment in its native habitat. Stephanie Helber, Gidon Winters, Marleen Stuhr, E. F. Belshe, Stefanie Bröhl, Michael Schmid, Hauke Reuter and Mirta Teichberg (2021). Frontiers in Plant Science.

 

36. Elective affinities or random choice within the seagrass holobiont? The case of the native Posidonia oceanica (L.) Delile and the exotic Halophila stipulacea (Forssk.) Asch. from the same site (Limassol, Cyprus). Chiara Conte, Alice Rotini, Gidon Winters, Marlen I. Vasquez, Giulia Piazza, Demetris Kletou, Luciana Migliore (2021). Aquatic Botany 174: 103420.

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35. Temporal and Spatial Changes in Phyllosphere Microbiome of Acacia Trees Growing in Arid Environments. Ashraf Al Ashhab, Shiri Meshner, Rivka Alexander-Shani, Hana Dimerets, Michael Brandwein, Yael Bar-Lavan and Gidon Winters (2021). Frontiers in Microbiology 12:656269.

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34. Oil Pollution Affects the Central Metabolism of Keystone Vachellia (Acacia) Trees. Marco Ferrante, Anuma Dangol, Shoshana Didi-Cohen, Gidon Winters, Vered Tzin, Michal Segoli (2021). Sustainability 13 (12), 6660.

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33. Unexpectedly low δ 13C in leaves, branches, stems and roots of three acacia species growing in hyper-arid environments. Daphna Uni, Elli Groner, Elaine Soloway, Amgad Hjazin, Spencer Johnswick, Gidon Winters, Efrat Sheffer, Ido Rog, Yael Wagner, Tamir Klein (2021). Journal of Plant Ecology 14 (1): 117-131.

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32. The Seagrass Holobiont: What We Know and What We Still Need to Disclose for Its Possible Use as an Ecological Indicator. Conte, Chiara; Rotini, Alice; Manfra, Loredana; D'Andrea, Marco M.; Winters, Gidon; Migliore, Luciana (2021). Water 13(4): 406.

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31. Projected Rapid Habitat Expansion of Tropical Seagrass Species in the Mediterranean Sea as Climate Change Progresses. P Beca-Carretero, M Teichberg, G Winters, G Procaccini, H Reuter (2020). Frontiers in Plant Science 11:1762.

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30. The tropical seagrass Halophila stipulacea: reviewing what we know from its native and invasive habitats, alongside identifying knowledge gaps. Winters, G., et al. (2020). Frontiers in Marine Science 6: 812.

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29. Responses of the seagrass Halophila stipulacea to depth and spatial gradients in its native region (Red Sea): morphology, in situ growth and biomass production. Azcárate-García T., Beca-Carretero P., Villamayor B., Stengel D.B., Winters G. (2020). Aquatic Botany 165: 103252.

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28. Responses of invasive and native populations of the seagrass Halophila stipulacea to simulated climate change. Manh Nguyen H., Yadav N.S., Barak S., Lima F.P., Sapir Y., Winters G. (2020). Frontiers in Marine Science 6:812.

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27. Seasonal dynamics of native and invasive Halophila stipulacea populations – A case study from the northern Gulf of Aqaba and the eastern Mediterranean Sea. Manh Nguyen H., Savva I., Kleitou P., Kletou D., Lima F.P., Sapir Y., Winters G. (2020). Aquatic Botany 162, 103205.

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26. Halophila stipulacea descriptors in the native area (Red Sea): a baseline for future comparisons with native and non-native populations. Beca-Carretero, Alice Rotini, Astrid Mejia, Luciana Migliore, Salvatrice Vizzini and Gidon Winters (2020). Marine Environmental Research 153, 104828.

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25. Depth-induced adjustment of fatty acid and pigment composition suggests high biochemical plasticity in the tropical seagrass Halophila stipulacea. Beca-Carretero P., Guihéneuf F., Winters G., Stengel D.B. (2019). Marine Ecology Progress Series 608:105-117.

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24. Tree growth and water-use in hyper-arid Acacia occurs during the hottest and driest season. Winters G., Otieno D., Cohen S., Bogner C., Ragowloski G., Paudel I., Klein T. (2018). Oecologia 188: 695-705.

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23. Differences in flowering sex ratios between native and invasive populations of the seagrass Halophila stipulacea. Manh Nguyen H., Kleitou P., Kletou D., Sapir Y., Winters G. (2018). Botanica Marina 61: 337-342.

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22. The tropical invasive seagrass, Halophila stipulacea, has a superior ability to tolerate dynamic changes in salinity levels compared to its freshwater relative, Vallisneria Americana. Oscar M.A., Barak S., Winters G. (2018). Frontiers in Plant Science 9:950.

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21. Detecting hierarchical levels of connectivity in a population of Acacia tortilis at the northern edge of the species' global distribution: Combining classical population genetics and network analyses. Rodger Y.S., Greenbaum G., Silver M., Bar-David S., Winters G. (2018). PLoS ONE 13:e0194901.

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20. Germination, physiological and biochemical responses of acacia seedlings (Acacia raddiana and Acacia tortilis) to petroleum contaminated soils. Hoai Tran T., Mayzlish-Gati E., Eshel A., Winters G. (2018). Environmental Pollution 234: 642-655.

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19. Physiological and biogeochemical responses of super-corals to thermal stress from the northern Gulf of Aqaba, Red Sea. Grottoli A.G, Tchernov D., Winters G. (2017). Frontiers in Marine Science 4.

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18. Advancing marine conservation in European and contiguous seas with the MarCons Action. Katsanevakis S., et al. (2017). Research Ideas and Outcomes 3: e11884.

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17. A standardized protocol to monitor Acacia trees in the Arava. Groner E., et al. (2017). Negev, Dead Sea and Arava Studies 9: 1-14.

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16. Ecophysiological plasticity and bacteriome shift in the seagrass Halophila stipulacea along a depth

gradient in the Northern Red Sea. Rotini A., Mejia A. Y., Costa R., Migliore L., Winters G. (2017). Frontiers in Plant Science. 2016:2015.

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15. A low-cost field-survey method for mapping seagrasses and their potential threats: an example from the northern Gulf of Aqaba, Red Sea. Winters G., Edelist D., Shem-Tov R., Beer S., Rilov G. (2017). Aquatic Conservation: Marine and Freshwater Ecosystems 27: 324-339.

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14. Assessing the ecological status of seagrasses using morphology, biochemical descriptors and microbial community analyses. A study in Halophila stipulacea meadows in the northern Red Sea. Mejia A.Y., Rotini A., Lacasella F., Bookman R., Thaller M.C., Shem-Tov R., Winters G., Migliore L. (2016). Ecological Indicators 60: 1150-1163.

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13. Mapping underground layers in the super arid Gidron Wadi using electrical resistivity tomography (ERT). Winters G., Ryvkin I., Rudkov T., Moreno Z., Furman A. (2015). Journal of Arid Environments 121: 79-83.

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12. Genome-wide transcriptomic responses of the seagrasses Zostera marina and Nanozostera noltii under a simulated heatwave confirm functional types. Franssen U.S., et al. (2014). Marine Genomics 15:65-73.

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11. Characterizations of microsatellite DNA markers for Acacia tortilis. Winters G., Shklar G., Korol L. (2013). Conservation Genetics Resources 5: 807-809.

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10. Identifying core features of adaptive metabolic mechanisms for chronic heat stress attenuation contributing to systems robustness. Gu J., et al. (2012). Integrative Biology 4: 480-493.

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9. Transcriptomic resilience to global warming in the seagrass Zostera marina, a marine foundation species. Franssen U.S., et al. (2011). Proceedings of the National Academy of Sciences 108: 19276-19281.

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8. The effects of a simulated heat wave on the photophysiology and gene expression of high and low-latitude populations of Zostera marina. Winters G., Nelle P., Fricke B., Reusch T.B.H. (2011). Marine Ecology Progress Series 435: 83-95.

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7. Population-specificity of heat stress gene induction in northern and southern eelgrass Zostera marina populations under simulated global warming. Bergmann N., Winters G., Rauch G., Eizaguirre C., Gu J., Nelle P., Fricke B., Reusch T.B.H. (2010). Molecular Ecology 19: 2870-2883.

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6. Photographic assessment of coral chlorophyll contents: implications for ecophysiological studies and coral monitoring. Winters G., Holzman R., Blekhman A., Beer S., Loya Y. (2009b). Journal of Experimental Marine Biology and Ecology 380: 25-35.

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5. Spatial and temporal photoacclimation of Stylophora pistillata: zooxanthella size, pigmentation, location and clade. Winters G., Beer S., Ben Zvi B., Z., Brickner I., Loya Y. (2009a). Marine Ecology Progress Series 384: 107-119.

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4. Cellular pathology and histopathology of hypo-salinity exposure on the coral Stylophora pistillata. Downs C.A., Kramarsky-Winter E., Woodley C.M., Downs A., Winters G., Loya Y., Ostrander G.K. (2009). Science of the Total Environment 407: 4838-4851.

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3. Adaptation strategies of the corallimorpharian Rhodactis rhodostoma to irradiance and temperature. Kuguru B., Winters G., Santos S.R., Beer S., Chadwick N.E. (2007). Marine Biology 151:1287-1298.

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2. In situ measured seasonal variations in Fv/Fm of two common Red Sea corals. Winters G., Loya Y., Beer S. (2006). Coral Reefs 25: 593-598.

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1. Photoinhibition in shallow-water colonies of the coral Stylophora pistillata as measured in situ. Winters G., Loya Y., Routtgers R., Beer S. (2003). Limnology and Oceanography 48: 1388-1393.

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